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DNA-directed_RNA_polymerase_subunit_alpha

Euk-Vir

Marseillevirus_marseillevirus

DNA-directed_RNA_polymerase_subunit_alpha__YP_003406803__Marseillevirus_marseillevirus__694581

Identity

Accession:
YP_003406803 ↗
Protein ID:
DNA-directed_RNA_polymerase_subunit_alpha
Kingdom:
euk

Quality

71.1 mean pLDDT

3D Structure

Domains

high = three domain segmentations agree, medium = two domain segmentations agree. Numbered high-first then medium, N→C within each tier.

D1 medium residues 138-202_231-242
PDB
Domain cluster: representative
CATH (5)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
1j20A02 3.90.1260.10 Alpha Beta › Alpha-Beta Complex › Argininosuccinate synthetase, chain A, domain 2 › Argininosuccinate synthetase, chain A, domain 2 0.57 41.0 3.03e-01 76.6% 95.8%
4r8oA00 2.40.128.720 Mainly Beta › Beta Barrel › Lipocalin › 0.53 36.0 3.42e-01 72.7% 90.8%
3kxeA00 3.30.2310.20 Alpha Beta › 2-Layer Sandwich › YaeB-like fold › RelE-like 0.51 32.0 3.02e-01 74.0% 48.9%
5cegD00 3.30.2310.20 Alpha Beta › 2-Layer Sandwich › YaeB-like fold › RelE-like 0.51 34.0 3.14e-01 88.3% 51.5%
4nhxA01 2.60.120.620 Mainly Beta › Sandwich › Jelly Rolls › q2cbj1_9rhob like domain 0.50 38.0 2.86e-01 84.4% 65.8%
ECOD (4)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
5031337 375.8.1.1 few secondary structure elements › Rubredoxin-like › Zinc-binding domain of translation initiation factor 2 beta › Zinc-binding domain of translation initiation factor 2 beta › eIF-5_eIF-2B 0.64 28.0 3.70e-01 75.3% 74.4%
4381748 241.1.1.2 a+b two layers › Type III secretory system chaperone-like › Type III secretory system chaperone › Type III secretory system chaperone › Autophagy_act_C 0.54 46.0 3.46e-01 100.0% 87.9%
1291229 101.1.2.194 alpha arrays › HTH › HTH › winged helix domain › RIOX1_C_WH 0.52 37.0 3.13e-01 76.6% 72.7%
1712440 4312.1.1.3 a+b two layers › RelE-like › RelE-like › RelE-like › ParE_toxin 0.51 34.0 3.15e-01 88.3% 50.5%
D2 medium residues 272-293_307-365
PDB
Domain cluster: representative
CATH (13)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
2pvqA02 1.20.1050.10 Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › 0.69 45.0 4.07e-01 87.7% 50.9%
2yxhA00 1.10.287.1080 Mainly Alpha › Orthogonal Bundle › Helix Hairpins › MazG-like 0.65 39.0 3.51e-01 82.7% 41.6%
2imiB02 1.20.1050.10 Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › 0.65 43.0 3.59e-01 86.4% 40.0%
4mk3A02 1.20.1050.10 Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › 0.64 42.0 3.78e-01 90.1% 47.8%
4gltA02 1.20.1050.10 Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › 0.64 44.0 3.95e-01 86.4% 51.8%
8g0lB01 1.25.40.1040 Mainly Alpha › Alpha Horseshoe › Serine Threonine Protein Phosphatase 5, Tetratricopeptide repeat › 0.63 37.0 2.34e-01 72.8% 12.1%
3anwA01 1.20.58.1030 Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › 0.63 55.0 4.88e-01 95.1% 72.6%
5l0wB01 1.25.40.10 Mainly Alpha › Alpha Horseshoe › Serine Threonine Protein Phosphatase 5, Tetratricopeptide repeat › Tetratricopeptide repeat domain 0.63 36.0 3.06e-01 74.1% 32.6%
1p5zB00 3.40.50.300 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › P-loop containing nucleotide triphosphate hydrolases 0.62 41.0 2.96e-01 88.9% 23.6%
2e9xA01 1.20.58.1030 Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › 0.60 50.0 4.34e-01 92.6% 70.3%
3ug9A02 1.20.1070.10 Mainly Alpha › Up-down Bundle › Rhopdopsin 7-helix transmembrane proteins › Rhodopsin 7-helix transmembrane proteins 0.60 42.0 3.04e-01 98.8% 26.2%
3sqiA01 1.10.150.540 Mainly Alpha › Orthogonal Bundle › DNA polymerase; domain 1 › 0.53 31.0 2.92e-01 74.1% 44.4%
1i51B00 3.30.70.1470 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Caspase-like 0.51 32.0 3.09e-01 90.1% 55.4%
ECOD (15)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
3533047 109.4.1.1324 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › P4Ha_N, TPR_P4H 0.73 39.0 2.83e-01 80.2% 21.0%
4933335 604.12.1.1 alpha bundles › Spectrin repeat-like › MIT domain › MIT domain › MIT 0.71 45.0 4.60e-01 91.4% 66.3%
3960865 604.12.1.21 alpha bundles › Spectrin repeat-like › MIT domain › MIT domain › DUF3017 0.70 41.0 4.12e-01 80.2% 56.5%
3203288 101.1.1.0 alpha arrays › HTH › HTH › Three-helical HTH 0.69 39.0 4.21e-01 85.2% 64.3%
3615590 603.1.1.0 alpha bundles › STAT-like › t-snare proteins › t-snare proteins 0.65 46.0 4.11e-01 97.5% 50.8%
3290838 192.29.1.0 alpha bundles › Long alpha-hairpin › bMERB domain (bivalent Mical/EHBP Rab binding) › bMERB domain (bivalent Mical/EHBP Rab binding) 0.64 39.0 3.44e-01 75.3% 42.6%
3827442 605.1.1.0 alpha duplicates or obligate multimers › ROP-like › Homodimeric domain of signal transducing histidine kinase › Homodimeric domain of signal transducing histidine kinase 0.63 37.0 3.83e-01 88.9% 61.3%
3727265 2004.1.1.30 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › Helicase_C 0.62 46.0 3.16e-01 76.5% 37.8%
293479 109.1.1.7 alpha superhelices › Repetitive alpha hairpins › Glutathione S-transferase (GST)-C › Glutathione S-transferase (GST)-C › GST_C_3 0.61 36.0 3.20e-01 87.7% 40.0%
3193797 109.24.1.0 alpha superhelices › Repetitive alpha hairpins › Helical domain in dedicator of cytokinesis protein 9 › Helical domain in dedicator of cytokinesis protein 9 0.59 43.0 3.73e-01 86.4% 51.7%
3592536 2004.1.1.0 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases 0.57 41.0 3.02e-01 76.5% 28.4%
4001599 109.3.1.2 alpha superhelices › Repetitive alpha hairpins › Ankyrin repeat › Ankyrin repeat › Ank,Ank_2 0.54 34.0 2.42e-01 81.5% 19.2%
3532664 3755.3.1.391 alpha bundles › YscO-like › CT398 helical hairpin › CT398 helical hairpin › USHBP1_PDZ-bd 0.52 43.0 3.57e-01 90.1% 62.1%
3208854 2004.1.1.430 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › ABC_tran, ABC_tran_Xtn 0.51 44.0 2.62e-01 97.5% 41.6%
3280303 3684.1.1.0 alpha complex topology › PSPTO4464 C-terminal domain-like › PSPTO4464 C-terminal domain-like › PSPTO4464 C-terminal domain-like 0.50 44.0 3.84e-01 100.0% 88.0%
D3 medium residues 367-487
PDB
Pfam (1)
AccessionNameScoreE-valueQ covHMM cov
PF00623.26 best RNA_pol_Rpb1_2 61.4 1.80e-16 100.0% 75.9%