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IMGVR_UViG_3300004640_000048-3300004640-Ga0066615_12522824

Arc-Vir

IMGVR_UViG_3300004640_000048-3300004640-Ga0066615_12522824

Quality

87.9 mean pLDDT

Cluster

Singleton — not in a non-trivial cluster

3D Structure

Domains

high = three domain segmentations agree, medium = two domain segmentations agree. Numbered high-first then medium, N→C within each tier.

D1 high residues 8-48
PDB
Domain cluster: representative
CATH (59)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
1wv4B00 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.76 59.0 4.03e-01 100.0% 23.4%
2hq7B00 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.73 56.0 3.86e-01 95.1% 24.6%
2i51B00 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.68 55.0 3.58e-01 100.0% 19.9%
4qqgG00 2.30.30.140 Mainly Beta › Roll › SH3 type barrels. › 0.68 54.0 4.64e-01 95.1% 54.2%
4h75A00 2.80.10.70 Mainly Beta › Trefoil › Trefoil (Acidic Fibroblast Growth Factor, subunit A) › Spindlin/Ssty 0.67 53.0 3.43e-01 95.1% 18.6%
5jv4A00 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.65 53.0 3.78e-01 97.6% 28.9%
3lxrF00 1.10.4120.20 Mainly Alpha › Orthogonal Bundle › SopE-like GEF fold › 0.65 43.0 2.87e-01 100.0% 16.0%
2fg9A01 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.64 48.0 3.33e-01 97.6% 21.4%
2kr7A02 2.40.10.330 Mainly Beta › Beta Barrel › Thrombin, subunit H › 0.64 44.0 4.09e-01 95.1% 54.4%
2v1yA00 3.10.20.90 Alpha Beta › Roll › Ubiquitin-like (UB roll) › Phosphatidylinositol 3-kinase Catalytic Subunit; Chain A, domain 1 0.64 46.0 3.68e-01 80.5% 87.6%
5yjlD01 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.64 49.0 3.41e-01 97.6% 24.6%
4lduA03 2.30.30.1040 Mainly Beta › Roll › SH3 type barrels. › 0.63 48.0 4.30e-01 95.1% 70.6%
3ne5B01 2.40.420.20 Mainly Beta › Beta Barrel › conserved putative lor/sdh protein from methanococcus maripaludis s2 fold › 0.63 46.0 3.82e-01 95.1% 40.7%
4hh3A01 3.30.450.20 Alpha Beta › 2-Layer Sandwich › Beta-Lactamase › PAS domain 0.62 47.0 3.56e-01 100.0% 32.5%
1vl7A00 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.61 46.0 3.32e-01 100.0% 26.7%
2byvE05 3.10.20.90 Alpha Beta › Roll › Ubiquitin-like (UB roll) › Phosphatidylinositol 3-kinase Catalytic Subunit; Chain A, domain 1 0.61 47.0 3.86e-01 87.8% 86.1%
1xfsA00 3.30.530.20 Alpha Beta › 2-Layer Sandwich › Alpha-D-Glucose-1,6-Bisphosphate; Chain A, domain 4 › START domain 0.60 46.0 3.12e-01 100.0% 22.7%
4qrlA00 2.40.128.280 Mainly Beta › Beta Barrel › Lipocalin › 0.60 44.0 3.40e-01 87.8% 63.6%
1hwyA02 3.40.50.10860 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Leucine Dehydrogenase, chain A, domain 1 0.59 43.0 3.03e-01 82.9% 87.2%
5ay6A01 2.60.98.20 Mainly Beta › Sandwich › Tick-borne Encephalitis virus Glycoprotein; domain 1 › Flagellar hook protein FlgE 0.59 43.0 2.94e-01 82.9% 73.1%
2rceA02 2.40.10.10 Mainly Beta › Beta Barrel › Thrombin, subunit H › Trypsin-like serine proteases 0.59 47.0 3.56e-01 100.0% 35.8%
3gasB02 2.30.110.10 Mainly Beta › Roll › Pnp Oxidase; Chain A › Electron Transport, Fmn-binding Protein; Chain A 0.58 42.0 2.92e-01 97.6% 20.2%
1v1qA00 2.40.50.140 Mainly Beta › Beta Barrel › OB fold (Dihydrolipoamide Acetyltransferase, E2P) › Nucleic acid-binding proteins 0.58 45.0 3.56e-01 100.0% 85.5%
1g5hA01 3.30.930.10 Alpha Beta › 2-Layer Sandwich › BirA Bifunctional Protein; domain 2 › Bira Bifunctional Protein; Domain 2 0.58 47.0 2.91e-01 100.0% 70.2%
7c5wA01 3.10.450.50 Alpha Beta › Roll › Nuclear Transport Factor 2; Chain: A, › 0.58 49.0 3.37e-01 97.6% 33.1%
5hmaA02 2.40.10.10 Mainly Beta › Beta Barrel › Thrombin, subunit H › Trypsin-like serine proteases 0.57 47.0 3.66e-01 100.0% 42.3%
3iiiA01 3.40.50.1820 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Alpha/Beta hydrolase fold, catalytic domain 0.57 49.0 2.99e-01 100.0% 33.2%
3pijA01 2.115.10.20 Mainly Beta › 5 Propeller › Tachylectin-2; Chain A › Glycosyl hydrolase domain; family 43 0.57 47.0 2.73e-01 97.6% 11.0%
2v4dE01 2.40.420.20 Mainly Beta › Beta Barrel › conserved putative lor/sdh protein from methanococcus maripaludis s2 fold › 0.56 41.0 3.53e-01 97.6% 42.9%
3k7cA00 3.10.450.50 Alpha Beta › Roll › Nuclear Transport Factor 2; Chain: A, › 0.56 47.0 3.43e-01 97.6% 35.2%
3fgqA01 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.55 45.0 3.01e-01 97.6% 48.4%
4ic5A02 2.40.10.10 Mainly Beta › Beta Barrel › Thrombin, subunit H › Trypsin-like serine proteases 0.55 45.0 3.51e-01 100.0% 43.3%
1nqzA00 3.90.79.10 Alpha Beta › Alpha-Beta Complex › Nucleoside Triphosphate Pyrophosphohydrolase › Nucleoside Triphosphate Pyrophosphohydrolase 0.55 40.0 2.73e-01 100.0% 19.9%
2peeB02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.55 41.0 3.07e-01 92.7% 80.2%
1imvA01 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.54 44.0 3.03e-01 100.0% 55.6%
1hp7A01 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.54 44.0 3.51e-01 100.0% 95.8%
3er7B00 3.10.450.50 Alpha Beta › Roll › Nuclear Transport Factor 2; Chain: A, › 0.54 44.0 3.24e-01 97.6% 41.5%
1zj8A04 3.30.413.10 Alpha Beta › 2-Layer Sandwich › Sulfite Reductase Hemoprotein; domain 1 › Sulfite Reductase Hemoprotein, domain 1 0.54 44.0 3.10e-01 100.0% 39.2%
1olzA01 2.130.10.10 Mainly Beta › 7 Propeller › Methylamine Dehydrogenase; Chain H › YVTN repeat-like/Quinoprotein amine dehydrogenase 0.54 44.0 2.53e-01 100.0% 17.3%
4bq2D01 2.60.120.430 Mainly Beta › Sandwich › Jelly Rolls › Galactose-binding lectin 0.54 43.0 2.80e-01 90.2% 50.8%
3le2A02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.54 39.0 2.98e-01 92.7% 79.4%
7jooC01 2.60.40.10 Mainly Beta › Sandwich › Immunoglobulin-like › Immunoglobulins 0.53 40.0 3.07e-01 82.9% 69.1%
3njaA02 2.10.70.100 Mainly Beta › Ribbon › Complement Module; domain 1 › 0.53 42.0 4.12e-01 100.0% 84.8%
2v95A02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.53 42.0 3.12e-01 100.0% 95.3%
1f0cA02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.53 42.0 3.20e-01 100.0% 80.5%
2vlgC00 3.30.450.20 Alpha Beta › 2-Layer Sandwich › Beta-Lactamase › PAS domain 0.53 43.0 3.36e-01 100.0% 40.2%
1h7zA00 2.60.90.10 Mainly Beta › Sandwich › Adenovirus Type 5 Fiber Protein (Receptor Binding Domain) › Adenovirus pIV-related, attachment domain 0.52 45.0 2.89e-01 100.0% 22.0%
3f02B02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.52 41.0 3.28e-01 100.0% 97.0%
3icyA00 3.30.450.20 Alpha Beta › 2-Layer Sandwich › Beta-Lactamase › PAS domain 0.52 43.0 3.20e-01 100.0% 96.6%
5inwA02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.52 41.0 3.18e-01 97.6% 85.2%
5iu1B00 3.30.450.20 Alpha Beta › 2-Layer Sandwich › Beta-Lactamase › PAS domain 0.52 40.0 3.07e-01 95.1% 34.8%
5ncsA02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.52 39.0 2.84e-01 90.2% 79.9%
1uzxA00 3.10.110.10 Alpha Beta › Roll › Ubiquitin Conjugating Enzyme › Ubiquitin Conjugating Enzyme 0.51 42.0 3.05e-01 100.0% 47.9%
3mfxB00 3.30.450.20 Alpha Beta › 2-Layer Sandwich › Beta-Lactamase › PAS domain 0.51 36.0 2.86e-01 95.1% 31.6%
6mlyB01 2.115.10.20 Mainly Beta › 5 Propeller › Tachylectin-2; Chain A › Glycosyl hydrolase domain; family 43 0.51 39.0 2.46e-01 92.7% 92.0%
3vpyA00 2.60.200.20 Mainly Beta › Sandwich › Tumour Suppressor Smad4 › 0.51 40.0 2.91e-01 100.0% 49.7%
4zk3A02 2.30.39.10 Mainly Beta › Roll › Alpha-1-antitrypsin; domain 1 › Alpha-1-antitrypsin, domain 1 0.50 40.0 2.97e-01 100.0% 79.5%
1nltA03 2.60.260.20 Mainly Beta › Sandwich › HSP40/DNAj peptide-binding domain › Urease metallochaperone UreE, N-terminal domain 0.50 36.0 3.08e-01 85.4% 68.3%
2il5A00 3.30.530.20 Alpha Beta › 2-Layer Sandwich › Alpha-D-Glucose-1,6-Bisphosphate; Chain A, domain 4 › START domain 0.50 39.0 2.72e-01 95.1% 85.2%
ECOD (69)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
3614175 148.1.3.0 alpha arrays › Histone-like › Histone-related › AAA+ ATPase lid domain 0.76 58.0 4.28e-01 95.1% 31.8%
3713334 4.8.1.0 beta barrels › SH3 › Chromo domain-like › Chromo domain-like 0.75 61.0 5.49e-01 95.1% 65.0%
3704395 4.1.1.0 beta barrels › SH3 › SH3 › SH3 0.71 56.0 5.04e-01 95.1% 60.9%
3812766 4.1.1.42 beta barrels › SH3 › SH3 › SH3 › Agenet 0.71 56.0 4.87e-01 95.1% 58.6%
2978978 4.1.1.42 beta barrels › SH3 › SH3 › SH3 › Agenet 0.71 59.0 4.89e-01 95.1% 53.3%
3679362 4.1.1.351 beta barrels › SH3 › SH3 › SH3 › SH3_ISE2 0.71 61.0 4.84e-01 100.0% 78.8%
4986455 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.69 54.0 3.34e-01 97.6% 13.8%
3608562 4.1.1.0 beta barrels › SH3 › SH3 › SH3 0.68 53.0 3.66e-01 95.1% 23.9%
4929587 1.1.13.0 beta barrels › cradle loop barrel › RIFT-related › Phage tail proteins 0.67 55.0 4.43e-01 100.0% 50.0%
3501223 7516.1.1.8 a/b three-layered sandwiches › Nucleotide-diphospho-sugar transferases › Nucleotide-diphospho-sugar transferases › Nucleotide-diphospho-sugar transferases › Galactosyl_T 0.67 52.0 3.16e-01 87.8% 72.1%
3594429 4.6.1.0 beta barrels › SH3 › PRC-barrel domain › PRC-barrel domain 0.67 53.0 4.15e-01 95.1% 53.5%
3447254 1.1.5.33 beta barrels › cradle loop barrel › RIFT-related › FMN-binding split barrel › Trypsin_2 0.66 53.0 3.37e-01 100.0% 17.3%
3946167 1.1.7.80 beta barrels › cradle loop barrel › RIFT-related › Alanine racemase-C › RND-MFP_C 0.66 49.0 3.88e-01 95.1% 36.8%
3969410 1.1.1.0 beta barrels › cradle loop barrel › RIFT-related › acid protease 0.65 52.0 3.92e-01 100.0% 42.0%
4512383 1.1.7.0 beta barrels › cradle loop barrel › RIFT-related › Alanine racemase-C 0.64 51.0 4.01e-01 95.1% 46.0%
3938317 389.1.1.0 few secondary structure elements › EGF-like › EGF-related › EGF/Laminin 0.64 43.0 4.38e-01 97.6% 72.5%
3340613 4.1.1.0 beta barrels › SH3 › SH3 › SH3 0.63 51.0 4.26e-01 100.0% 50.0%
4387111 4.1.1.97 beta barrels › SH3 › SH3 › SH3 › DUF150_C 0.63 50.0 4.15e-01 95.1% 48.8%
3894729 4.1.1.461 beta barrels › SH3 › SH3 › SH3 › zf-CCCH 0.61 49.0 3.99e-01 100.0% 43.3%
3212698 9.1.1.0 beta barrels › Lipocalins/Streptavidin › Lipocalins › Lipocalins 0.61 51.0 3.40e-01 100.0% 71.1%
4960095 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.61 43.0 3.09e-01 78.0% 52.3%
3685401 4.1.1.20 beta barrels › SH3 › SH3 › SH3 › BAH 0.61 49.0 3.01e-01 97.6% 26.3%
4933932 304.139.1.1 a+b two layers › Alpha-beta plaits › Cas7-related › CRISPR-associated protein Cas7/Csa2-related › DevR 0.60 46.0 2.84e-01 90.2% 78.4%
3608755 4106.1.1.1 few secondary structure elements › Zinc hairpin stack › Zinc hairpin stack › Zinc hairpin stack › DHHC 0.60 44.0 3.69e-01 87.8% 60.0%
4114383 4.8.1.47 beta barrels › SH3 › Chromo domain-like › Chromo domain-like › zf_CCCH_4 0.60 48.0 3.90e-01 100.0% 45.9%
4530545 4.1.1.217 beta barrels › SH3 › SH3 › SH3 › zf-CCCH_4 0.59 48.0 3.82e-01 100.0% 41.1%
4973551 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.59 49.0 3.53e-01 95.1% 31.2%
4235293 4.1.1.0 beta barrels › SH3 › SH3 › SH3 0.59 45.0 3.51e-01 95.1% 41.7%
4170378 2.1.1.6 beta barrels › OB-fold › Nucleic acid-binding protein › Nucleic acid-binding protein › SSB 0.59 49.0 3.63e-01 97.6% 70.4%
154136 4059.1.1.0 a+b complex topology › Serpins › Serpins › Serpins 0.58 48.0 2.80e-01 100.0% 25.6%
3608162 216.1.1.0 a+b two layers › UBC-like › UBC-like › UBC-like 0.58 45.0 3.09e-01 90.2% 83.6%
5048407 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.58 46.0 3.27e-01 97.6% 29.2%
4940002 223.1.1.122 a+b three layers › Profilin-like › sensor domains › sensor domains › HisKA 0.58 45.0 2.94e-01 97.6% 19.0%
5047587 223.1.1.2 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS 0.58 47.0 3.41e-01 95.1% 31.2%
4945264 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.58 45.0 3.40e-01 97.6% 34.5%
4997160 223.1.1.25 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_9 0.57 44.0 3.47e-01 95.1% 38.0%
3772106 1.1.17.3 beta barrels › cradle loop barrel › RIFT-related › FMN-binding split barrel 2 › Trypsin_2 0.57 47.0 3.01e-01 100.0% 18.7%
4939999 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.57 43.0 3.34e-01 97.6% 34.5%
3056322 1.1.7.7 beta barrels › cradle loop barrel › RIFT-related › Alanine racemase-C › FAD_binding_6 0.56 43.0 3.21e-01 87.8% 73.6%
3634173 216.1.1.3 a+b two layers › UBC-like › UBC-like › UBC-like › UEV 0.56 45.0 3.13e-01 97.6% 69.7%
3940272 2498.1.1.14 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › Peptidase_M8 0.56 43.0 2.58e-01 100.0% 47.9%
5082808 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.56 45.0 2.44e-01 100.0% 4.2%
4980684 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.55 44.0 2.53e-01 97.6% 8.6%
3620976 10.4.1.0 beta sandwiches › jelly-roll › Spermadhesin, CUB domain › Spermadhesin, CUB domain 0.55 42.0 2.68e-01 80.5% 20.0%
4989092 223.1.1.25 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_9 0.55 40.0 3.11e-01 95.1% 33.3%
5004858 223.1.1.25 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_9 0.54 46.0 3.21e-01 100.0% 27.3%
4959595 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.54 42.0 2.62e-01 97.6% 14.6%
4999616 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.54 42.0 2.53e-01 95.1% 10.1%
5004657 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.54 40.0 2.58e-01 100.0% 14.9%
5008034 223.1.1.25 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_9 0.53 44.0 3.16e-01 95.1% 31.2%
4634739 389.1.1.0 few secondary structure elements › EGF-like › EGF-related › EGF/Laminin 0.53 38.0 3.64e-01 100.0% 62.0%
5061315 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.53 42.0 2.59e-01 100.0% 12.9%
4959633 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.53 41.0 2.59e-01 95.1% 16.1%
3255683 223.1.1.24 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_8 0.52 40.0 2.93e-01 95.1% 28.1%
5021847 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.52 43.0 2.71e-01 95.1% 15.9%
137752 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.52 43.0 3.20e-01 100.0% 96.6%
3443030 330.1.1.0 a+b two layers › dsRBD-like › dsRNA-binding domain-like › dsRNA-binding domain-like 0.52 39.0 3.31e-01 100.0% 46.3%
4959266 223.1.1.2 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS 0.52 41.0 2.60e-01 100.0% 15.5%
4551342 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.52 44.0 3.46e-01 95.1% 45.9%
3192325 376.1.1.43 few secondary structure elements › RING/U-box-like › RING/U-box-like › RING/U-box › PRT6_C 0.51 40.0 2.71e-01 97.6% 79.4%
397995 223.1.1.13 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_3 0.51 39.0 3.04e-01 95.1% 36.1%
5061316 223.1.1.14 a+b three layers › Profilin-like › sensor domains › sensor domains › PAS_4 0.51 39.0 2.87e-01 100.0% 29.2%
3460576 109.3.1.162 alpha superhelices › Repetitive alpha hairpins › Ankyrin repeat › Ankyrin repeat › Ank, Ank_2, Ank_5 0.51 43.0 2.67e-01 100.0% 31.9%
3634800 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.51 41.0 3.16e-01 97.6% 38.1%
3994642 389.2.1.0 few secondary structure elements › EGF-like › Serine protease inhibitors › Serine protease inhibitors 0.51 34.0 3.53e-01 95.1% 77.1%
5011151 319.1.1.1 beta sandwiches › HSP20-like › HSP20-like chaperones › HSP20-like chaperones › HSP20 0.51 43.0 2.89e-01 100.0% 25.0%
3974170 4.1.1.0 beta barrels › SH3 › SH3 › SH3 0.50 38.0 3.17e-01 100.0% 58.0%
3933168 9.1.1.0 beta barrels › Lipocalins/Streptavidin › Lipocalins › Lipocalins 0.50 44.0 2.92e-01 100.0% 70.0%
4980710 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.50 38.0 2.96e-01 95.1% 33.9%