Back to structures

KY821088.1__ARM70158.1__HARAMBE_9__00009

Bact-Vir

KY821088.1__ARM70158.1__HARAMBE_9__00009

Identity

Accession:
KY821088 ↗
Kingdom:
phage

Quality

86.4 mean pLDDT

Cluster

Singleton — not in a non-trivial cluster

3D Structure

Domains

high = three domain segmentations agree, medium = two domain segmentations agree. Numbered high-first then medium, N→C within each tier.

D1 high residues 6-44
PDB
Domain cluster: representative
CATH (41)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
4grhA00 3.60.120.10 Alpha Beta › 4-Layer Sandwich › Anthranilate synthase › Anthranilate synthase 0.76 55.0 3.08e-01 76.9% 9.8%
7asgA02 2.30.180.10 Mainly Beta › Roll › FAS1 domain › FAS1 domain 0.71 49.0 3.51e-01 74.4% 73.1%
1egaA01 3.40.50.300 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › P-loop containing nucleotide triphosphate hydrolases 0.69 48.0 3.07e-01 74.4% 31.5%
2p62A01 3.40.50.10620 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › PH0156-like domains 0.68 46.0 3.11e-01 71.8% 42.6%
3gg6A00 3.90.79.10 Alpha Beta › Alpha-Beta Complex › Nucleoside Triphosphate Pyrophosphohydrolase › Nucleoside Triphosphate Pyrophosphohydrolase 0.66 53.0 3.67e-01 94.9% 59.0%
3lv0A01 3.30.540.10 Alpha Beta › 2-Layer Sandwich › Fructose-1,6-Bisphosphatase; Chain A, domain 1 › Fructose-1,6-Bisphosphatase, subunit A, domain 1 0.66 49.0 3.42e-01 84.6% 41.7%
3gr5A01 3.55.50.30 Alpha Beta › 3-Layer(bab) Sandwich › Phage tail protein beta-alpha-beta fold › 0.66 53.0 4.30e-01 100.0% 67.1%
2p3nA01 3.30.540.10 Alpha Beta › 2-Layer Sandwich › Fructose-1,6-Bisphosphatase; Chain A, domain 1 › Fructose-1,6-Bisphosphatase, subunit A, domain 1 0.65 49.0 3.41e-01 84.6% 40.7%
1f6bB00 3.40.50.300 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › P-loop containing nucleotide triphosphate hydrolases 0.63 44.0 2.84e-01 74.4% 25.0%
4qxdA01 3.30.540.10 Alpha Beta › 2-Layer Sandwich › Fructose-1,6-Bisphosphatase; Chain A, domain 1 › Fructose-1,6-Bisphosphatase, subunit A, domain 1 0.62 45.0 3.09e-01 82.1% 50.6%
3p26A01 3.40.50.300 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › P-loop containing nucleotide triphosphate hydrolases 0.60 47.0 2.93e-01 87.2% 30.2%
1n6zA00 3.10.20.250 Alpha Beta › Roll › Ubiquitin-like (UB roll) › YML108W-like 0.60 44.0 3.31e-01 84.6% 81.9%
1s5jA02 3.30.420.10 Alpha Beta › 2-Layer Sandwich › Nucleotidyltransferase; domain 5 › Ribonuclease H-like superfamily/Ribonuclease H 0.59 41.0 2.62e-01 76.9% 14.9%
1gxsB02 3.40.50.11320 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › 0.59 42.0 3.16e-01 76.9% 63.3%
1g12A00 3.40.390.10 Alpha Beta › 3-Layer(aba) Sandwich › Collagenase (Catalytic Domain) › Collagenase (Catalytic Domain) 0.59 47.0 3.21e-01 100.0% 52.1%
1uqtA01 3.40.50.2000 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Glycogen Phosphorylase B; 0.58 40.0 2.45e-01 74.4% 13.2%
4kmaA02 3.30.1360.230 Alpha Beta › 2-Layer Sandwich › Gyrase A; domain 2 › Sufu, C-terminal domain 0.57 48.0 3.39e-01 94.9% 57.1%
6p8iA04 2.70.130.10 Mainly Beta › Distorted Sandwich › Cation-dependent Mannose-6-phosphate Receptor; Chain A › Mannose-6-phosphate receptor binding domain 0.56 44.0 3.46e-01 100.0% 65.4%
2qy6A01 3.40.50.150 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Vaccinia Virus protein VP39 0.56 44.0 2.70e-01 87.2% 39.6%
2j4xA02 2.40.50.110 Mainly Beta › Beta Barrel › OB fold (Dihydrolipoamide Acetyltransferase, E2P) › 0.56 49.0 4.01e-01 100.0% 86.1%
5nkkF01 3.40.50.300 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › P-loop containing nucleotide triphosphate hydrolases 0.56 48.0 2.96e-01 100.0% 28.2%
6j7xA01 1.25.40.120 Mainly Alpha › Alpha Horseshoe › Serine Threonine Protein Phosphatase 5, Tetratricopeptide repeat › Protein prenylyltransferase 0.55 45.0 2.79e-01 100.0% 14.9%
2epkX01 3.30.160.230 Alpha Beta › 2-Layer Sandwich › Double Stranded RNA Binding Domain › N-acetyl-beta-d-glucosaminidase 0.55 43.0 3.59e-01 97.4% 47.5%
1qvvA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.55 44.0 2.83e-01 100.0% 79.9%
1pm3A00 2.30.30.240 Mainly Beta › Roll › SH3 type barrels. › PRC-barrel domain 0.55 44.0 3.68e-01 92.3% 89.9%
4p5pA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.54 46.0 2.91e-01 100.0% 79.1%
1u9cA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.54 45.0 2.86e-01 100.0% 77.8%
4bkwA02 3.30.1360.220 Alpha Beta › 2-Layer Sandwich › Gyrase A; domain 2 › Domain of unknown function (DUF3480), N-terminal subdomain 0.54 46.0 3.45e-01 100.0% 53.0%
8a3pA01 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.54 45.0 2.96e-01 100.0% 82.1%
2w2gB01 3.40.220.30 Alpha Beta › 3-Layer(aba) Sandwich › Leucine Aminopeptidase, subunit E; domain 1 › Nsp3, SUD-N subdomain 0.54 45.0 3.13e-01 94.9% 52.8%
4bl8B03 3.30.1360.230 Alpha Beta › 2-Layer Sandwich › Gyrase A; domain 2 › Sufu, C-terminal domain 0.54 47.0 3.22e-01 100.0% 50.0%
1sy7A03 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.53 38.0 2.56e-01 84.6% 36.4%
4gdhA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.52 44.0 2.90e-01 100.0% 74.7%
1e8cA01 3.40.1390.10 Alpha Beta › 3-Layer(aba) Sandwich › Udp-n-acetylmuramoylalanyl-d-glutamate--2,6- Diaminopimelate Ligase; Chain: A, domain 1 › MurE/MurF, N-terminal domain 0.52 45.0 3.39e-01 100.0% 82.2%
3ewnA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.52 43.0 2.70e-01 100.0% 55.0%
2jh3A01 3.40.50.1400 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › 0.52 43.0 2.93e-01 94.9% 78.9%
4mtnA02 2.40.50.140 Mainly Beta › Beta Barrel › OB fold (Dihydrolipoamide Acetyltransferase, E2P) › Nucleic acid-binding proteins 0.51 44.0 3.81e-01 100.0% 93.7%
1ac5A00 3.40.50.1820 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Alpha/Beta hydrolase fold, catalytic domain 0.51 36.0 2.10e-01 84.6% 12.4%
4y1eA00 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.51 43.0 2.87e-01 100.0% 79.5%
3fseA01 3.40.50.880 Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Class I glutamine amidotransferase (GATase) domain 0.51 42.0 2.77e-01 100.0% 68.5%
1ipaA02 3.40.1280.10 Alpha Beta › 3-Layer(aba) Sandwich › Alpha/beta knot › SPOUT methyltransferase, trefoil knot domain 0.51 41.0 2.79e-01 100.0% 24.2%
ECOD (65)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
3782948 221.15.1.0 a+b two layers › beta-Grasp › beta-grasp fold domain in leucine-tRNA ligase › beta-grasp fold domain in leucine-tRNA ligase 0.77 54.0 4.70e-01 74.4% 100.0%
4503474 239.3.1.1 beta barrels › Ribosomal protein L25-like › FAS1 domain › FAS1 domain › Fasciclin 0.77 53.0 3.54e-01 74.4% 73.5%
5020150 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.73 54.0 4.16e-01 79.5% 65.9%
4960926 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.73 54.0 4.69e-01 79.5% 95.0%
3366512 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.70 46.0 2.79e-01 74.4% 10.2%
4989862 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.70 52.0 4.28e-01 79.5% 81.4%
4984657 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.70 52.0 4.05e-01 79.5% 67.1%
4062727 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.70 46.0 2.85e-01 74.4% 11.8%
4954187 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.68 50.0 4.40e-01 79.5% 95.0%
4952589 221.10.1.1 a+b two layers › beta-Grasp › FdhD/NarQ beta-grasp fold domain › FdhD/NarQ beta-grasp fold domain › FdhD-NarQ 0.68 49.0 4.34e-01 79.5% 95.0%
3601536 2004.1.1.0 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases 0.68 47.0 2.99e-01 76.9% 17.0%
4420884 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.66 44.0 2.81e-01 74.4% 13.0%
4184406 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.66 43.0 2.67e-01 74.4% 10.2%
3261838 63.1.1.0 beta barrels › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain 0.66 55.0 3.83e-01 100.0% 67.1%
3263895 63.1.1.0 beta barrels › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain 0.65 53.0 3.69e-01 100.0% 75.3%
4989777 5.1.4.0 beta duplicates or obligate multimers › beta-propeller-like › beta-propeller › 7-bladed 0.65 48.0 2.87e-01 82.1% 26.3%
3284640 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.65 43.0 2.62e-01 74.4% 9.3%
4049237 63.1.1.5 beta barrels › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › ATG27 0.65 52.0 3.64e-01 100.0% 70.0%
3337596 2004.1.1.150 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › DRMBL 0.64 54.0 3.71e-01 100.0% 89.0%
4429065 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.64 42.0 2.62e-01 74.4% 10.6%
3622893 207.1.1.0 beta duplicates or obligate multimers › Single-stranded right-handed beta-helix › Leucine-rich repeats › Leucine-rich repeats 0.64 50.0 2.90e-01 87.2% 21.4%
3256855 2004.1.1.150 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › DRMBL 0.63 53.0 3.62e-01 100.0% 81.3%
3944717 2004.1.1.0 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases 0.63 53.0 3.18e-01 92.3% 15.2%
3208455 211.1.1.0 a+b two layers › Glyoxalase/Bleomycin resistance protein/Dihydroxybiphenyl dioxygenase › Glyoxalase/Bleomycin resistance protein/Dihydroxybiphenyl dioxygenase › Glyoxalase/Bleomycin resistance protein/Dihydroxybiphenyl dioxygenase 0.63 49.0 4.41e-01 87.2% 81.5%
4969796 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.62 49.0 3.71e-01 100.0% 36.0%
4221242 63.1.1.5 beta barrels › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › ATG27 0.62 52.0 3.57e-01 100.0% 70.7%
4979795 4.6.1.2 beta barrels › SH3 › PRC-barrel domain › PRC-barrel domain › PRC 0.61 46.0 3.55e-01 82.1% 82.2%
4165654 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.61 40.0 2.52e-01 74.4% 10.9%
5051240 375.1.1.0 few secondary structure elements › Rubredoxin-like › Rubredoxin-related › Rubredoxin-related 0.61 51.0 4.10e-01 94.9% 49.3%
4940509 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.60 50.0 3.77e-01 97.4% 40.0%
3205039 63.1.1.5 beta barrels › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › Mannose 6-phosphate receptor domain › ATG27 0.59 49.0 3.34e-01 100.0% 57.5%
3290645 2498.1.1.29 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › Peptidase_M78 0.59 52.0 3.67e-01 100.0% 32.5%
3954373 223.1.1.0 a+b three layers › Profilin-like › sensor domains › sensor domains 0.59 50.0 2.99e-01 100.0% 13.9%
3218082 145.1.1.1 alpha arrays › F-box domain › F-box domain › F-box domain › F-box 0.59 46.0 3.59e-01 89.7% 75.6%
3452884 109.3.1.239 alpha superhelices › Repetitive alpha hairpins › Ankyrin repeat › Ankyrin repeat › DUF247 0.59 45.0 3.10e-01 89.7% 23.8%
3380869 109.4.1.450 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › DUF247 0.58 44.0 2.56e-01 84.6% 9.3%
3421392 109.4.1.450 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › DUF247 0.58 45.0 2.65e-01 100.0% 11.1%
4199652 4.6.1.2 beta barrels › SH3 › PRC-barrel domain › PRC-barrel domain › PRC 0.58 47.0 3.90e-01 92.3% 90.0%
3458998 109.58.1.2 alpha superhelices › Repetitive alpha hairpins › DNA repair protein Rev1 C-terminal domain › DNA repair protein Rev1 C-terminal domain › DUF247 0.58 43.0 2.92e-01 84.6% 21.9%
3953551 2498.1.1.29 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › Peptidase_M78 0.57 44.0 2.81e-01 87.2% 34.5%
4950270 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.57 48.0 3.10e-01 100.0% 32.8%
3252321 148.1.3.0 alpha arrays › Histone-like › Histone-related › AAA+ ATPase lid domain 0.56 47.0 3.20e-01 97.4% 67.1%
4974865 7512.1.1.30 a/b three-layered sandwiches › UDP-Glycosyltransferase/glycogen phosphorylase › UDP-Glycosyltransferase/glycogen phosphorylase › UDP-Glycosyltransferase/glycogen phosphorylase › Glyco_transf_4 0.56 39.0 2.41e-01 74.4% 12.9%
4990363 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.56 38.0 2.44e-01 71.8% 15.8%
1122687 2007.1.1.0 a/b three-layered sandwiches › Flavodoxin-like › Class I glutamine amidotransferase-like › Class I glutamine amidotransferase-like 0.56 47.0 2.83e-01 100.0% 64.8%
3989086 2498.1.1.29 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › Peptidase_M78 0.56 47.0 3.05e-01 100.0% 22.1%
3962296 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.55 47.0 2.96e-01 100.0% 18.3%
5027859 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.55 47.0 3.24e-01 100.0% 37.7%
3839024 4070.1.1.2 alpha arrays › FtsH protease domain-like › FtsH protease domain-like › FtsH protease domain-like › Peptidase_M50 0.55 40.0 2.55e-01 82.1% 26.7%
3471910 4018.1.1.2 a+b two layers › a+b domain in carbohydrate phosphatases › a+b domain in carbohydrate phosphatases › a+b domain in carbohydrate phosphatases › Inositol_P 0.54 46.0 3.07e-01 100.0% 54.1%
4310284 2498.1.1.143 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › DUF3318 0.54 44.0 2.83e-01 100.0% 18.5%
5016974 7584.1.1.0 a/b three-layered sandwiches › Rossmann-like domain in Acetyl-CoA synthetase-like proteins › Rossmann-like domain in Acetyl-CoA synthetase-like proteins › Rossmann-like domain in Acetyl-CoA synthetase-like proteins 0.53 45.0 2.67e-01 100.0% 46.1%
3427656 109.4.1.450 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › DUF247 0.53 42.0 2.56e-01 94.9% 12.4%
4014518 261.1.1.0 a+b complex topology › Hect, E3 ligase catalytic domain › Hect, E3 ligase catalytic domain › Hect, E3 ligase catalytic domain 0.53 45.0 3.08e-01 100.0% 76.0%
3399410 2.6.1.0 beta barrels › OB-fold › Staphylococcal nuclease › Staphylococcal nuclease 0.53 43.0 3.57e-01 100.0% 83.7%
3470915 109.4.1.18 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › PPTA 0.53 46.0 2.68e-01 100.0% 11.0%
5024540 2498.1.1.0 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" 0.53 44.0 3.07e-01 100.0% 26.9%
3494153 109.4.1.18 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › PPTA 0.53 45.0 2.84e-01 100.0% 18.1%
4928334 4262.1.1.1 a/b three-layered sandwiches › CobE/CbiG C-terminal domain-like › CobE/CbiG C-terminal domain-like › CobE/CbiG C-terminal domain-like › CbiG_C 0.53 41.0 2.79e-01 100.0% 23.2%
4959341 2008.1.1.152 a/b three-layered sandwiches › Restriction endonuclease-like › Restriction endonuclease-like › Restriction endonuclease-like › DUF2130 0.53 41.0 3.10e-01 100.0% 36.0%
3180442 109.4.1.18 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › PPTA 0.52 41.0 2.48e-01 100.0% 12.4%
3936153 2498.1.1.39 mixed a+b and a/b › Zincin-like › "Metalloproteases (""zincins"") catalytic domain" › "Metalloproteases (""zincins"") catalytic domain" › SprT-like 0.52 43.0 3.00e-01 100.0% 30.0%
3390173 109.4.1.18 alpha superhelices › Repetitive alpha hairpins › ARM repeat › ARM repeat › PPTA 0.52 41.0 2.46e-01 100.0% 10.5%
5038258 2004.1.1.0 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases 0.51 40.0 2.64e-01 87.2% 42.6%
4013028 170.2.1.0 alpha bundles › Retrovirus capsid protein › Retrovirus capsid protein N-terminal domain › Retrovirus capsid protein N-terminal domain 0.51 36.0 2.61e-01 76.9% 27.8%