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RNA-dependent_RNA_polymerase
Euk-VirHubei_narna-like_virus_21
RNA-dependent_RNA_polymerase__YP_009337775__Hubei_narna-like_virus_21__1922952
Identity
- Accession:
- YP_009337775 ↗
- Protein ID:
- RNA-dependent_RNA_polymerase
- Kingdom:
- euk
Quality
67.8
mean pLDDT
Cluster
Singleton — not in a non-trivial cluster
3D Structure
Domains
high = three domain segmentations agree, medium = two domain segmentations agree. Numbered high-first then medium, N→C within each tier.
D1
medium
residues 1-98
Domain cluster:
representative
CATH (16)
| Domain ID | Class ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 3iqcA00 | 1.20.120.340 | Mainly Alpha › Up-down Bundle › Four Helix Bundle (Hemerythrin (Met), subunit A) › Flagellar protein FliS | 0.64 | 39.0 | 3.69e-01 | 82.7% | 50.4% |
| 3lnnB03 | 1.10.287.470 | Mainly Alpha › Orthogonal Bundle › Helix Hairpins › Helix hairpin bin | 0.61 | 39.0 | 4.57e-01 | 79.6% | 94.0% |
| 3i01A01 | 1.20.1270.30 | Mainly Alpha › Up-down Bundle › Substrate Binding Domain Of Dnak; Chain:A; Domain 2 › | 0.61 | 45.0 | 3.66e-01 | 77.6% | 62.1% |
| 1aluA00 | 1.20.1250.10 | Mainly Alpha › Up-down Bundle › Growth Hormone; Chain: A; › | 0.60 | 36.0 | 3.09e-01 | 81.6% | 36.9% |
| 3qx3A04 | 1.10.268.10 | Mainly Alpha › Orthogonal Bundle › Topoisomerase; domain 3 › Topoisomerase, domain 3 | 0.60 | 43.0 | 4.27e-01 | 73.5% | 99.0% |
| 1abvA00 | 1.10.520.20 | Mainly Alpha › Orthogonal Bundle › Peroxidase; domain 1 › N-terminal domain of the delta subunit of the F1F0-ATP synthase | 0.60 | 42.0 | 4.18e-01 | 73.5% | 82.9% |
| 2d9dA00 | 1.20.58.120 | Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › BAG domain | 0.58 | 36.0 | 3.78e-01 | 81.6% | 67.4% |
| 6lccA01 | 1.10.600.10 | Mainly Alpha › Orthogonal Bundle › Farnesyl Diphosphate Synthase › Farnesyl Diphosphate Synthase | 0.58 | 45.0 | 3.05e-01 | 83.7% | 25.8% |
| 2m6bA00 | 1.20.58.390 | Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › Neurotransmitter-gated ion-channel transmembrane domain | 0.57 | 42.0 | 3.71e-01 | 78.6% | 98.0% |
| 1jqkA03 | 1.20.1270.30 | Mainly Alpha › Up-down Bundle › Substrate Binding Domain Of Dnak; Chain:A; Domain 2 › | 0.57 | 41.0 | 3.41e-01 | 75.5% | 63.4% |
| 1hm7B00 | 1.10.600.10 | Mainly Alpha › Orthogonal Bundle › Farnesyl Diphosphate Synthase › Farnesyl Diphosphate Synthase | 0.57 | 45.0 | 3.20e-01 | 85.7% | 31.1% |
| 3d85C00 | 1.20.1250.10 | Mainly Alpha › Up-down Bundle › Growth Hormone; Chain: A; › | 0.56 | 41.0 | 3.74e-01 | 76.5% | 99.2% |
| 6kd7A00 | 1.10.600.10 | Mainly Alpha › Orthogonal Bundle › Farnesyl Diphosphate Synthase › Farnesyl Diphosphate Synthase | 0.55 | 46.0 | 3.16e-01 | 88.8% | 55.7% |
| 1aepA00 | 1.20.120.20 | Mainly Alpha › Up-down Bundle › Four Helix Bundle (Hemerythrin (Met), subunit A) › Apolipoprotein | 0.55 | 37.0 | 3.20e-01 | 85.7% | 43.8% |
| 2pbeA02 | 1.20.120.330 | Mainly Alpha › Up-down Bundle › Four Helix Bundle (Hemerythrin (Met), subunit A) › Nucleotidyltransferases domain 2 | 0.54 | 38.0 | 3.53e-01 | 73.5% | 84.9% |
| 4hz4A02 | 1.20.1050.10 | Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › | 0.50 | 44.0 | 4.20e-01 | 94.9% | 92.9% |
ECOD (11)
| UID | F-ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 5058148 | 601.4.1.0 ↗ | alpha bundles › Four-helical up-and-down bundle › Histidine kinase associated sensor domains › Histidine kinase associated sensor domains | 0.63 | 48.0 | 4.77e-01 | 78.6% | 99.0% |
| 3282529 | 632.1.1.3 ↗ | alpha bundles › immunoglobulin/albumin-binding domain-like › Families 57/38 glycoside transferase middle domain › Families 57/38 glycoside transferase middle domain › Alpha-mann_mid | 0.58 | 34.0 | 3.35e-01 | 79.6% | 53.3% |
| 3441715 | 603.1.1.0 ↗ | alpha bundles › STAT-like › t-snare proteins › t-snare proteins | 0.55 | 43.0 | 4.13e-01 | 84.7% | 72.7% |
| 3962961 | 150.5.1.53 ↗ | alpha bundles › Ferritin/Heme oxygenase/4-helical cytokines › ESAT-6 like › ESAT-6 like › PPE | 0.55 | 42.0 | 4.61e-01 | 78.6% | 96.2% |
| 3957966 | 150.8.1.1 ↗ | alpha bundles › Ferritin/Heme oxygenase/4-helical cytokines › PPE › PPE › PPE | 0.54 | 41.0 | 3.87e-01 | 78.6% | 67.0% |
| 4017135 | 603.1.1.0 ↗ | alpha bundles › STAT-like › t-snare proteins › t-snare proteins | 0.52 | 30.0 | 3.07e-01 | 81.6% | 58.9% |
| 3954841 | 150.5.1.11 ↗ | alpha bundles › Ferritin/Heme oxygenase/4-helical cytokines › ESAT-6 like › ESAT-6 like › T7SS_ESX_EspC | 0.52 | 40.0 | 4.00e-01 | 79.6% | 79.0% |
| 3979729 | 3755.3.1.500 ↗ | alpha bundles › YscO-like › CT398 helical hairpin › CT398 helical hairpin › IpaC_SipC | 0.52 | 41.0 | 3.79e-01 | 82.7% | 80.0% |
| 3284573 | 150.5.1.11 ↗ | alpha bundles › Ferritin/Heme oxygenase/4-helical cytokines › ESAT-6 like › ESAT-6 like › T7SS_ESX_EspC | 0.51 | 39.0 | 3.92e-01 | 79.6% | 79.0% |
| 3781441 | 3291.1.1.0 ↗ | alpha bundles › Charged multivesicular body protein 3 (CHMP3)-related › Charged multivesicular body protein 3 (CHMP3)-related › Charged multivesicular body protein 3 (CHMP3)-related | 0.51 | 39.0 | 3.97e-01 | 79.6% | 89.5% |
| 5014397 | 5050.1.1.9 ↗ | alpha complex topology › Major facilitator superfamily (MFS) general substrate transporter › Major facilitator superfamily (MFS) general substrate transporter › Major facilitator superfamily (MFS) general substrate transporter › MFS_1 | 0.50 | 38.0 | 3.13e-01 | 81.6% | 64.2% |
D2
medium
residues 99-109_287-311_349-380
Domain cluster:
representative
CATH (10)
| Domain ID | Class ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 4epzA00 | 1.25.40.810 | Mainly Alpha › Alpha Horseshoe › Serine Threonine Protein Phosphatase 5, Tetratricopeptide repeat › UpxZ | 0.64 | 39.0 | 2.97e-01 | 100.0% | 27.3% |
| 1ct5A00 | 3.20.20.10 | Alpha Beta › Alpha-Beta Barrel › TIM Barrel › Alanine racemase | 0.58 | 42.0 | 2.92e-01 | 76.5% | 38.2% |
| 1gq2A02 | 3.40.50.720 | Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › NAD(P)-binding Rossmann-like Domain | 0.55 | 39.0 | 2.68e-01 | 79.4% | 85.3% |
| 3r2qA02 | 1.20.1050.10 | Mainly Alpha › Up-down Bundle › Glutathione S-transferase Yfyf (Class Pi); Chain A, domain 2 › | 0.55 | 36.0 | 3.08e-01 | 100.0% | 40.7% |
| 2zj2A04 | 1.10.3380.20 | Mainly Alpha › Orthogonal Bundle › Sec63 N-terminal domain-like fold › | 0.55 | 44.0 | 3.31e-01 | 88.2% | 39.5% |
| 5ix8A02 | 3.40.50.2300 | Alpha Beta › 3-Layer(aba) Sandwich › Rossmann fold › Response regulator | 0.54 | 43.0 | 3.39e-01 | 91.2% | 95.0% |
| 3anwB00 | 1.20.58.2050 | Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › | 0.53 | 40.0 | 3.06e-01 | 82.4% | 81.5% |
| 1zoyA03 | 1.20.58.100 | Mainly Alpha › Up-down Bundle › Methane Monooxygenase Hydroxylase; Chain G, domain 1 › Fumarate reductase/succinate dehydrogenase flavoprotein-like, C-terminal domain | 0.51 | 38.0 | 3.29e-01 | 82.4% | 77.8% |
| 3p01A01 | 6.10.140.590 | Special › Helix non-globular › Helix Hairpins › | 0.51 | 39.0 | 3.61e-01 | 98.5% | 66.3% |
| 3ajfA00 | 1.20.1440.190 | Mainly Alpha › Up-down Bundle › de novo design (two linked rop proteins) › Tenuivirus movement protein | 0.50 | 37.0 | 3.38e-01 | 79.4% | 64.1% |
ECOD (6)
| UID | F-ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 3684651 | 181.1.1.1 ↗ | alpha bundles › Domain of the SRP/SRP receptor G proteins-like › Domain of the SRP/SRP receptor G-proteins › Domain of the SRP/SRP receptor G-proteins › SRP54_N | 0.63 | 36.0 | 3.37e-01 | 73.5% | 47.1% |
| 5049338 | 2007.1.4.2 ↗ | a/b three-layered sandwiches › Flavodoxin-like › Class I glutamine amidotransferase-like › Phosphofructokinase N-terminal domain › DAGK_cat | 0.56 | 42.0 | 3.44e-01 | 79.4% | 60.0% |
| 3677130 | 103.1.1.34 ↗ | alpha arrays › RuvA-C › RuvA-C, UBA, CRAL/TRIO-N, HBS1 › UBA-like domain › PIR2-like_helical | 0.54 | 41.0 | 3.33e-01 | 82.4% | 92.6% |
| 3343812 | 7073.1.1.1 ↗ | alpha arrays › C-terminal domain of Malic enzyme › C-terminal domain of Malic enzyme › C-terminal domain of Malic enzyme › Malic_M | 0.51 | 36.0 | 3.14e-01 | 100.0% | 48.6% |
| 3638639 | 1128.1.1.1 ↗ | alpha bundles › LYR protein › LYR protein › LYR protein › Complex1_LYR | 0.51 | 39.0 | 3.43e-01 | 82.4% | 68.0% |
| 3214991 | 101.1.2.10 ↗ | alpha arrays › HTH › HTH › winged helix domain › Linker_histone | 0.51 | 40.0 | 3.79e-01 | 100.0% | 70.6% |
D3
medium
residues 110-165_177-251_312-348
Domain cluster:
representative
CATH (1)
| Domain ID | Class ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 5mmjo00 | 1.10.287.10 | Mainly Alpha › Orthogonal Bundle › Helix Hairpins › S15/NS1, RNA-binding | 0.54 | 21.0 | 3.11e-01 | 72.6% | 78.7% |
ECOD (14)
| UID | F-ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 2850105 | 304.48.1.19 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › Bunya_RdRp | 0.63 | 46.0 | 3.94e-01 | 75.6% | 64.9% |
| 4152428 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.60 | 53.0 | 4.12e-01 | 93.5% | 54.1% |
| 3589612 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.59 | 55.0 | 4.36e-01 | 99.4% | 60.0% |
| 4138932 | 304.48.1.72 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_N | 0.58 | 49.0 | 3.97e-01 | 88.7% | 62.3% |
| 3173834 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.58 | 53.0 | 3.93e-01 | 98.8% | 45.6% |
| 4004424 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.57 | 53.0 | 4.04e-01 | 99.4% | 52.0% |
| 4497954 | 304.48.1.73 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1, RVT_N | 0.57 | 53.0 | 4.26e-01 | 98.8% | 62.9% |
| 3945039 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.57 | 53.0 | 4.16e-01 | 99.4% | 57.0% |
| 4872037 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.57 | 47.0 | 3.99e-01 | 86.9% | 65.8% |
| 5029718 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.57 | 47.0 | 3.67e-01 | 86.9% | 54.7% |
| 4461237 | 4967.1.1.0 ↗ | alpha bundles › helical bundle domain in reverse transcriptase-like polymerases › helical bundle domain in reverse transcriptase-like polymerases › helical bundle domain in reverse transcriptase-like polymerases | 0.56 | 52.0 | 3.64e-01 | 99.4% | 36.3% |
| 5002351 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.56 | 51.0 | 3.89e-01 | 97.6% | 50.7% |
| 5018583 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.55 | 51.0 | 4.10e-01 | 98.8% | 59.0% |
| 5018572 | 304.48.1.72 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_N | 0.50 | 41.0 | 4.07e-01 | 86.3% | 84.4% |
D4
medium
residues 252-286_381-522
Domain cluster:
representative
CATH (11)
| Domain ID | Class ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 5cqgA04 | 3.30.70.2630 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › | 0.64 | 38.0 | 4.79e-01 | 88.1% | 97.2% |
| 4mz0B05 | 3.30.70.250 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Malonyl-CoA ACP transacylase, ACP-binding | 0.57 | 23.0 | 3.75e-01 | 77.4% | 100.0% |
| 1konA03 | 3.30.70.980 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › YebC, transcriptional regulation domain | 0.56 | 24.0 | 3.62e-01 | 78.5% | 93.3% |
| 1zpwX00 | 3.30.70.240 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › | 0.55 | 25.0 | 3.62e-01 | 81.9% | 92.7% |
| 1lfpA03 | 3.30.70.980 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › YebC, transcriptional regulation domain | 0.54 | 24.0 | 3.55e-01 | 78.5% | 98.6% |
| 1h3dA03 | 3.30.70.120 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › | 0.53 | 22.0 | 3.42e-01 | 77.4% | 100.0% |
| 2ewhA01 | 3.30.70.1710 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › BMC (bacterial microcompartment) domain | 0.52 | 26.0 | 3.73e-01 | 80.2% | 100.0% |
| 2iboA00 | 3.30.70.930 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › | 0.52 | 27.0 | 3.76e-01 | 80.2% | 100.0% |
| 3d68A01 | 3.30.70.340 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Metallocarboxypeptidase-like | 0.52 | 26.0 | 3.49e-01 | 81.9% | 91.3% |
| 1vk8A00 | 3.30.70.930 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › | 0.51 | 27.0 | 3.69e-01 | 80.2% | 98.9% |
| 1kwmA01 | 3.30.70.340 | Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Metallocarboxypeptidase-like | 0.51 | 25.0 | 3.43e-01 | 79.7% | 94.3% |
ECOD (16)
| UID | F-ID | Classification | TM-score | Bits | E-value | Q cov | T cov |
|---|---|---|---|---|---|---|---|
| 4944359 | 304.56.1.0 ↗ | a+b two layers › Alpha-beta plaits › CRISPR associated protein Cas2-like › CRISPR associated protein Cas2-like | 0.58 | 26.0 | 3.73e-01 | 80.8% | 89.4% |
| 4235667 | 304.56.1.2 ↗ | a+b two layers › Alpha-beta plaits › CRISPR associated protein Cas2-like › CRISPR associated protein Cas2-like › CRISPR_Cas2 | 0.57 | 26.0 | 3.74e-01 | 81.4% | 90.6% |
| 4507546 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.57 | 24.0 | 3.78e-01 | 76.3% | 100.0% |
| 4123628 | 304.56.1.2 ↗ | a+b two layers › Alpha-beta plaits › CRISPR associated protein Cas2-like › CRISPR associated protein Cas2-like › CRISPR_Cas2 | 0.57 | 26.0 | 3.81e-01 | 80.8% | 96.2% |
| 5007506 | 304.5.1.0 ↗ | a+b two layers › Alpha-beta plaits › GlnB-like › GlnB-like | 0.56 | 25.0 | 3.52e-01 | 78.0% | 85.9% |
| 4279811 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.56 | 24.0 | 3.49e-01 | 76.8% | 87.5% |
| 4502075 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.56 | 23.0 | 3.70e-01 | 80.2% | 100.0% |
| 4936417 | 304.26.1.3 ↗ | a+b two layers › Alpha-beta plaits › MTH1187/YkoF-like › MTH1187/YkoF-like › DUF2102 | 0.55 | 29.0 | 3.64e-01 | 83.1% | 83.8% |
| 4070496 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.55 | 24.0 | 3.58e-01 | 79.1% | 96.0% |
| 3395435 | 304.11.1.0 ↗ | a+b two layers › Alpha-beta plaits › Probable ACP-binding domain of malonyl-CoA ACP transacylase › Probable ACP-binding domain of malonyl-CoA ACP transacylase | 0.55 | 23.0 | 3.53e-01 | 76.3% | 100.0% |
| 4231349 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.54 | 24.0 | 3.55e-01 | 78.0% | 100.0% |
| 4295716 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.54 | 24.0 | 3.54e-01 | 77.4% | 100.0% |
| 5023503 | 304.110.1.0 ↗ | a+b two layers › Alpha-beta plaits › Acylphosphatase-like › Acylphosphatase-like | 0.53 | 23.0 | 3.45e-01 | 77.4% | 98.6% |
| 4156338 | 304.117.1.1 ↗ | a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg | 0.52 | 23.0 | 3.39e-01 | 78.5% | 96.0% |
| 3927365 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.51 | 39.0 | 3.50e-01 | 95.5% | 57.6% |
| 3643305 | 304.48.1.1 ↗ | a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 | 0.50 | 37.0 | 4.26e-01 | 84.7% | 100.0% |
D5
medium
residues 536-616