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RNA-dependent_RNA_polymerase

Euk-Vir

Gigaspora_margarita_mitovirus_4

RNA-dependent_RNA_polymerase__YP_009551961__Gigaspora_margarita_mitovirus_4__2082668

Identity

Accession:
YP_009551961 ↗
Protein ID:
RNA-dependent_RNA_polymerase
Kingdom:
euk

Quality

49.9 mean pLDDT

Cluster

Singleton — not in a non-trivial cluster

3D Structure

Domains

high = three domain segmentations agree, medium = two domain segmentations agree. Numbered high-first then medium, N→C within each tier.

D1 medium residues 26-100_148-172
PDB
D2 medium residues 173-296
PDB
CATH (10)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
2jbwA01 1.20.1440.110 Mainly Alpha › Up-down Bundle › de novo design (two linked rop proteins) › acylaminoacyl peptidase 0.63 41.0 4.41e-01 99.2% 77.7%
2x1dA02 1.10.10.2120 Mainly Alpha › Orthogonal Bundle › Arc Repressor Mutant, subunit A › 0.60 32.0 3.94e-01 71.0% 83.8%
3pivA00 1.20.1250.10 Mainly Alpha › Up-down Bundle › Growth Hormone; Chain: A; › 0.55 44.0 4.13e-01 86.3% 84.6%
4k84A00 1.10.3520.10 Mainly Alpha › Orthogonal Bundle › Glycolipid transfer protein, GLTP › Glycolipid transfer protein 0.55 49.0 4.18e-01 99.2% 90.8%
1tdpA00 1.20.1440.50 Mainly Alpha › Up-down Bundle › de novo design (two linked rop proteins) › Ta0600-like 0.54 39.0 4.09e-01 100.0% 83.8%
1f5qB02 1.10.472.10 Mainly Alpha › Orthogonal Bundle › Cyclin A; domain 1 › Cyclin-like 0.54 38.0 3.69e-01 91.9% 64.5%
4nt1A00 1.10.3520.10 Mainly Alpha › Orthogonal Bundle › Glycolipid transfer protein, GLTP › Glycolipid transfer protein 0.54 47.0 4.01e-01 95.2% 95.5%
2nq2A00 1.10.3470.10 Mainly Alpha › Orthogonal Bundle › ABC transporter involved in vitamin B12 uptake, BtuC › ABC transporter involved in vitamin B12 uptake, BtuC 0.53 42.0 3.14e-01 83.1% 90.9%
2vcnA01 1.10.520.10 Mainly Alpha › Orthogonal Bundle › Peroxidase; domain 1 › 0.53 42.0 4.02e-01 84.7% 75.4%
3wvoC02 1.10.132.100 Mainly Alpha › Orthogonal Bundle › Topoisomerase I; Chain A, domain 4 › 0.50 43.0 4.30e-01 98.4% 90.0%
ECOD (6)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
3809284 2004.1.1.30 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › Helicase_C 0.65 54.0 4.02e-01 100.0% 36.4%
3835030 604.5.1.7 alpha bundles › Spectrin repeat-like › PhoU-like (Pfam 01895) › PhoU-like (Pfam 01895) › ALMT 0.60 54.0 4.92e-01 100.0% 95.2%
4472717 3930.1.1.0 alpha bundles › Helical bundle insertion in helicase domains › Helical bundle in Hef helicase › Helical bundle in Hef helicase 0.59 48.0 5.12e-01 100.0% 99.1%
4110399 5050.1.1.9 alpha complex topology › Major facilitator superfamily (MFS) general substrate transporter › Major facilitator superfamily (MFS) general substrate transporter › Major facilitator superfamily (MFS) general substrate transporter › MFS_1 0.58 46.0 3.67e-01 83.9% 89.2%
3250971 616.1.1.0 alpha arrays › S15/NS1 RNA-binding domain › S15/NS1 RNA-binding domain › S15/NS1 RNA-binding domain 0.58 43.0 4.74e-01 86.3% 98.0%
1487362 3930.2.1.1 alpha bundles › Helical bundle insertion in helicase domains › Helical bundle in endoribonuclease Dicer › Helical bundle in endoribonuclease Dicer › Dicer_PBD 0.54 46.0 4.75e-01 100.0% 96.6%
D3 medium residues 505-534_557-654
PDB
Pfam (1)
AccessionNameScoreE-valueQ covHMM cov
PF05919.17 best Mitovir_RNA_pol 65.7 5.80e-18 76.6% 15.7%
D4 medium residues 535-556_655-711
PDB
Domain cluster: representative
Pfam (1)
AccessionNameScoreE-valueQ covHMM cov
PF05919.17 best Mitovir_RNA_pol 60.4 2.30e-16 81.0% 10.3%
CATH (22)
Domain IDClass IDClassificationTM-scoreBitsE-valueQ covT cov
7uinD01 3.30.70.270 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Reverse transcriptase/Diguanylate cyclase domain 0.81 73.0 5.67e-01 100.0% 90.4%
1ayeA01 3.30.70.340 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Metallocarboxypeptidase-like 0.66 54.0 5.04e-01 88.6% 85.9%
2bvfA03 3.40.462.20 Alpha Beta › 3-Layer(aba) Sandwich › Vanillyl-alcohol Oxidase; Chain A, domain 3 › 0.64 47.0 3.52e-01 79.7% 74.3%
3d68A01 3.30.70.340 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Metallocarboxypeptidase-like 0.63 51.0 4.84e-01 87.3% 92.4%
1vdhA01 3.30.70.1030 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Apc35880; domain 1 0.62 49.0 4.29e-01 86.1% 85.1%
1kwmA01 3.30.70.340 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › Metallocarboxypeptidase-like 0.61 50.0 4.84e-01 88.6% 97.7%
1vk8A00 3.30.70.930 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › 0.59 46.0 4.40e-01 84.8% 97.8%
2hg4D03 3.40.366.10 Alpha Beta › 3-Layer(aba) Sandwich › Malonyl-Coenzyme A Acyl Carrier Protein; domain 2 › Malonyl-Coenzyme A Acyl Carrier Protein, domain 2 0.58 48.0 3.24e-01 91.1% 51.0%
1zpwX00 3.30.70.240 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › 0.57 44.0 4.38e-01 83.5% 98.8%
2aj2A02 3.30.70.1300 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › VC0467-like domains 0.57 37.0 3.57e-01 72.2% 57.8%
6u9hF02 3.30.70.1150 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › ACT-like. Chain A, domain 2 0.56 44.0 4.46e-01 83.5% 97.4%
1earA02 3.30.70.790 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › UreE, C-terminal domain 0.55 40.0 4.18e-01 75.9% 92.8%
1lfpA03 3.30.70.980 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › YebC, transcriptional regulation domain 0.55 44.0 4.59e-01 88.6% 100.0%
1gmuA01 3.30.70.790 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › UreE, C-terminal domain 0.54 38.0 4.02e-01 73.4% 92.5%
1zd0A01 3.30.2380.10 Alpha Beta › 2-Layer Sandwich › PF0523-like › CGI121/TPRKB 0.53 42.0 3.61e-01 86.1% 100.0%
5ijgA01 3.40.640.10 Alpha Beta › 3-Layer(aba) Sandwich › Aspartate Aminotransferase; domain 2 › Type I PLP-dependent aspartate aminotransferase-like (Major domain) 0.53 42.0 3.07e-01 84.8% 90.6%
3hx9B00 3.30.70.100 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › 0.52 40.0 3.77e-01 83.5% 98.0%
1ycyA00 2.30.30.100 Mainly Beta › Roll › SH3 type barrels. › 0.52 27.0 2.95e-01 96.2% 58.1%
1hc7A01 3.30.930.10 Alpha Beta › 2-Layer Sandwich › BirA Bifunctional Protein; domain 2 › Bira Bifunctional Protein; Domain 2 0.51 45.0 3.16e-01 100.0% 62.5%
2kviA00 3.30.70.330 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › RRM (RNA recognition motif) domain 0.51 39.0 4.00e-01 83.5% 100.0%
1wr2A02 3.30.1490.20 Alpha Beta › 2-Layer Sandwich › Dna Ligase; domain 1 › ATP-grasp fold, A domain 0.50 35.0 3.60e-01 73.4% 76.9%
1vbkA01 3.30.70.1510 Alpha Beta › 2-Layer Sandwich › Alpha-Beta Plaits › THUMP domain-like 0.50 37.0 3.73e-01 81.0% 98.8%
ECOD (38)
UIDF-IDClassificationTM-scoreBitsE-valueQ covT cov
3810170 304.48.1.43 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › Mitovir_RNA_pol 0.93 89.0 6.53e-01 100.0% 93.3%
3306901 304.48.1.43 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › Mitovir_RNA_pol 0.93 89.0 6.70e-01 100.0% 91.5%
3336938 304.48.1.43 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › Mitovir_RNA_pol 0.92 87.0 5.87e-01 100.0% 65.6%
3068775 304.48.1.1 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 0.82 75.0 5.48e-01 100.0% 74.9%
3934202 304.48.1.1 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 0.80 72.0 5.84e-01 100.0% 97.9%
3257066 304.48.1.1 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 0.76 68.0 4.81e-01 100.0% 56.7%
3315278 304.48.1.1 a+b two layers › Alpha-beta plaits › Adenylyl and guanylyl cyclase catalytic domain-like › Adenylyl and guanylyl cyclase catalytic domain-like › RVT_1 0.73 57.0 4.94e-01 86.1% 83.2%
4988159 304.37.1.1 a+b two layers › Alpha-beta plaits › Sulfite reductase, domains 1 and 3 › Sulfite reductase, domains 1 and 3 › NIR_SIR_ferr 0.71 49.0 4.36e-01 72.2% 87.8%
2038566 304.8.1.22 a+b two layers › Alpha-beta plaits › ACT-like › ACT-like › AHAS-like_ACT 0.66 46.0 4.58e-01 72.2% 100.0%
3859590 386.1.1.248 few secondary structure elements › beta-beta-alpha zinc fingers › beta-beta-alpha zinc fingers › beta-beta-alpha zinc fingers › zf-C2H2_ZNF592 0.66 41.0 4.81e-01 83.5% 98.0%
3405197 304.7.1.1 a+b two layers › Alpha-beta plaits › Protease propeptides/inhibitors › Protease propeptides/inhibitors › Propep_M14 0.64 51.0 5.11e-01 84.8% 100.0%
4060507 304.117.1.1 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg 0.64 44.0 4.60e-01 70.9% 100.0%
5054705 304.7.1.0 a+b two layers › Alpha-beta plaits › Protease propeptides/inhibitors › Protease propeptides/inhibitors 0.63 48.0 4.80e-01 81.0% 90.0%
3946828 304.11.1.0 a+b two layers › Alpha-beta plaits › Probable ACP-binding domain of malonyl-CoA ACP transacylase › Probable ACP-binding domain of malonyl-CoA ACP transacylase 0.63 44.0 4.76e-01 73.4% 100.0%
5047263 304.24.1.3 a+b two layers › Alpha-beta plaits › EF-G C-terminal domain-like › EF-G C-terminal domain-like › EFG_III 0.62 43.0 4.39e-01 72.2% 100.0%
4965210 304.163.1.0 a+b two layers › Alpha-beta plaits › ATP-binding protein TM_1403 insertion domain › ATP-binding protein TM_1403 insertion domain 0.61 44.0 4.96e-01 84.8% 100.0%
3593586 304.3.1.0 a+b two layers › Alpha-beta plaits › HMA-related › HMA, heavy metal-associated domain 0.61 47.0 4.74e-01 83.5% 93.8%
4263279 304.11.1.0 a+b two layers › Alpha-beta plaits › Probable ACP-binding domain of malonyl-CoA ACP transacylase › Probable ACP-binding domain of malonyl-CoA ACP transacylase 0.60 48.0 4.71e-01 86.1% 92.9%
4565839 304.120.1.0 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in ThiI › Ferredoxin-like domain in ThiI 0.60 43.0 4.79e-01 79.7% 100.0%
3961146 304.11.1.1 a+b two layers › Alpha-beta plaits › Probable ACP-binding domain of malonyl-CoA ACP transacylase › Probable ACP-binding domain of malonyl-CoA ACP transacylase › Acyl_transf_1 0.60 49.0 4.18e-01 89.9% 63.6%
5022748 304.56.1.2 a+b two layers › Alpha-beta plaits › CRISPR associated protein Cas2-like › CRISPR associated protein Cas2-like › CRISPR_Cas2 0.59 45.0 4.32e-01 81.0% 100.0%
4943089 304.126.1.0 a+b two layers › Alpha-beta plaits › ferredoxin-like domain in vacuolar ATP synthase subunit C › ferredoxin-like domain in vacuolar ATP synthase subunit C 0.59 41.0 4.17e-01 74.7% 100.0%
4614439 304.117.1.1 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg 0.58 46.0 4.60e-01 86.1% 100.0%
4944520 304.8.1.0 a+b two layers › Alpha-beta plaits › ACT-like › ACT-like 0.57 47.0 3.42e-01 91.1% 61.7%
4329557 304.24.1.0 a+b two layers › Alpha-beta plaits › EF-G C-terminal domain-like › EF-G C-terminal domain-like 0.56 45.0 4.31e-01 86.1% 93.3%
4520709 304.117.1.1 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg 0.56 44.0 4.43e-01 86.1% 100.0%
3723383 304.3.1.1 a+b two layers › Alpha-beta plaits › HMA-related › HMA, heavy metal-associated domain › HMA 0.56 43.0 4.33e-01 84.8% 96.2%
4070496 304.117.1.1 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg 0.56 43.0 4.47e-01 86.1% 97.3%
1145775 3268.1.1.1 a+b two layers › N-terminal domain in xanthine dehydrogenase › N-terminal domain in xanthine dehydrogenase › N-terminal domain in xanthine dehydrogenase › XdhC_CoxI 0.54 41.0 4.05e-01 81.0% 100.0%
3502962 4.1.1.89 beta barrels › SH3 › SH3 › SH3 › SM-ATX 0.54 31.0 3.10e-01 98.7% 52.5%
4990783 2004.1.1.73 a/b three-layered sandwiches › P-loop domains-like › P-loop domains-related › P-loop containing nucleoside triphosphate hydrolases › MMR_HSR1 0.54 38.0 2.83e-01 78.5% 26.9%
4030243 304.24.1.0 a+b two layers › Alpha-beta plaits › EF-G C-terminal domain-like › EF-G C-terminal domain-like 0.54 42.0 3.85e-01 84.8% 79.0%
5293 304.22.1.1 a+b two layers › Alpha-beta plaits › Urease metallochaperone UreE, C-terminal domain › Urease metallochaperone UreE, C-terminal domain › UreE_C 0.53 39.0 4.16e-01 79.7% 92.6%
3945340 235.1.1.9 a+b complex topology › Lysozyme-like › Lysozyme-like › Lysozyme-like › SLT 0.53 38.0 3.08e-01 75.9% 76.7%
None 0.52 39.0 3.52e-01 82.3% 95.7%
4156338 304.117.1.1 a+b two layers › Alpha-beta plaits › Ferredoxin-like domain in YebC › Ferredoxin-like domain in YebC › Transcrip_reg 0.52 41.0 4.24e-01 88.6% 97.3%
4215974 304.22.1.1 a+b two layers › Alpha-beta plaits › Urease metallochaperone UreE, C-terminal domain › Urease metallochaperone UreE, C-terminal domain › UreE_C 0.51 40.0 4.16e-01 87.3% 97.1%
4598958 4113.1.1.0 beta barrels › VC0467-like › VC0467-like › VC0467-like 0.51 43.0 3.35e-01 94.9% 82.8%
D5 medium residues 712-751_779-796
PDB